ecological parameterabiotic factoraerobic conditionanaerobic conditionbalance of matterbio-availabilitybiodiversitybiogeochemical cyclebiological attributebiological productionbiomassbiotic factorclimaxecological abundanceecological balanceecological carrying capacityecological factorecological nichegenetic diversitygenetic variationland covernutrientspeciesvegetation coverecology